:root {
      --parchment: #f3e9cd;
      --parchment-deep: #ead9b0;
      --parchment-light: #fdf8e9;
      --ink: #1a1f17;
      --forest: #243025;
      --forest-deep: #19211b;
      --accent: #3d5b3e;
      --umber: #7a5a3a;
      --rule: #8a7a4a;
      --rule-soft: #bca97a;
      --display: "EB Garamond", "Cardo", Georgia, serif;
      --mono: "IBM Plex Mono", ui-monospace, monospace;
    }
    *, *::before, *::after { box-sizing: border-box; }
    html, body { margin: 0; padding: 0; background: var(--parchment); color: var(--ink); font-family: var(--display); }
    body { font-size: 18px; line-height: 1.55; }
    a { color: inherit; text-decoration: none; }
    button { font: inherit; cursor: pointer; }
    .mono-cap { font-family: var(--mono); font-size: 11px; letter-spacing: 0.22em; text-transform: uppercase; opacity: 0.7; }

    /* ─── nav (shared) ─── */
    .nav { border-top: 3px double var(--rule); border-bottom: 1px solid rgba(138, 122, 74, 0.4); background: var(--parchment); position: relative; z-index: 10; }
    .nav-inner { display: flex; align-items: center; justify-content: space-between; padding: 20px 48px; max-width: 1180px; margin: 0 auto; }
    .wordmark { display: flex; align-items: baseline; gap: 14px; }
    .wordmark .tag { font-family: var(--mono); font-size: 9px; letter-spacing: 0.18em; text-transform: uppercase; opacity: 0.6; }
    .nav-links { display: flex; gap: 28px; align-items: center; }
    .nav-links a { font-family: var(--display); font-variant-caps: all-small-caps; letter-spacing: 0.14em; font-size: 16px; color: var(--ink); padding-bottom: 2px; border-bottom: 2px solid transparent; }
    .nav-links a.active { color: var(--accent); border-bottom-color: var(--accent); }
    .nav-links a:hover { color: var(--accent); }
    .nav-item { position: relative; display: flex; align-items: center; }
    .nav-menu { position: absolute; top: 100%; left: 50%; transform: translateX(-50%) translateY(4px); background: var(--parchment-light); border: 1px solid var(--rule); box-shadow: 0 12px 30px rgba(36, 48, 37, 0.16); padding: 6px; min-width: 176px; display: flex; flex-direction: column; opacity: 0; visibility: hidden; pointer-events: none; transition: opacity 150ms ease, transform 150ms ease; z-index: 40; }
    .nav-menu::before { content: ""; position: absolute; top: -10px; left: 0; right: 0; height: 10px; }
    .nav-item:hover .nav-menu, .nav-item:focus-within .nav-menu { opacity: 1; visibility: visible; pointer-events: auto; transform: translateX(-50%) translateY(0); }
    .nav-menu a { font-family: var(--display); font-variant-caps: all-small-caps; letter-spacing: 0.12em; font-size: 16px; padding: 9px 16px; color: var(--ink); border-bottom: none; white-space: nowrap; }
    .nav-menu a.active { color: var(--accent); }
    .nav-menu a:hover { background: var(--parchment-deep); color: var(--accent); }

    /* ─── page shell ─── */
    .wrap { max-width: 980px; margin: 0 auto; padding: 0 48px; }

    /* masthead */
    .reg-head { padding: 56px 0 28px; text-align: center; border-bottom: 3px double var(--rule); }
    .reg-kicker { color: var(--umber); }
    .reg-title { font-family: var(--display); font-weight: 500; font-size: 56px; line-height: 1.05; letter-spacing: -0.015em; margin: 14px 0 16px; color: var(--forest); }
    .reg-title em { font-style: italic; }
    .reg-lede { font-family: var(--display); font-style: italic; font-size: 21px; color: var(--umber); margin: 0 auto; max-width: 660px; line-height: 1.4; }
    .reg-divider { display: flex; align-items: center; gap: 14px; color: var(--rule); margin: 22px auto; max-width: 460px; }
    .reg-divider .rule { flex: 1; height: 1px; background: currentColor; opacity: 0.5; }
    .reg-divider .ornament { font-size: 20px; opacity: 0.7; line-height: 1; }

    /* search to change the focused entity */
    .reg-search { display: flex; gap: 0; max-width: 520px; margin: 0 auto; border: 1px solid var(--rule); background: var(--parchment-light); }
    .reg-search input { flex: 1; border: none; background: transparent; padding: 13px 16px; font-family: var(--display); font-size: 18px; color: var(--ink); outline: none; }
    .reg-search input::placeholder { color: rgba(26, 31, 23, 0.5); }
    .reg-search button { font-family: var(--display); font-variant-caps: all-small-caps; letter-spacing: 0.12em; font-size: 16px; background: var(--forest); color: var(--parchment); border: none; padding: 0 22px; }
    .reg-search button:hover { background: var(--forest-deep); }
    .reg-msg { min-height: 1.3em; margin: 12px 0 0; color: var(--umber); }
    .reg-chips { margin-top: 16px; display: flex; gap: 10px; justify-content: center; flex-wrap: wrap; align-items: baseline; }
    .reg-chips .chip { font-family: var(--display); font-style: italic; font-size: 16px; color: var(--forest); background: transparent; border: none; border-bottom: 1px dotted var(--rule); padding: 0 2px 2px; }
    .reg-chips .chip:hover { color: var(--accent); border-bottom-style: solid; }

    /* focused entity title block */
    .subject { padding: 40px 0 8px; }
    .subject-top { display: flex; gap: 44px; align-items: flex-start; justify-content: space-between; }
    .subject-text { flex: 1 1 auto; min-width: 0; }
    /* specimen plate: a Wikimedia photo (plants) or a PubChem 2D structure (compounds) */
    .subject-media { flex: 0 0 auto; width: 272px; margin-top: 4px; }
    .subject-media .plate {
      position: relative; width: 100%; height: 272px; overflow: hidden;
      background: #fdfaf1; border: 1px solid var(--rule);
      box-shadow: inset 0 0 0 1px rgba(122, 90, 58, 0.30), inset 0 0 0 4px #fdfaf1, inset 0 0 0 5px rgba(122, 90, 58, 0.14);
      display: flex; align-items: center; justify-content: center;
    }
    .subject-media img { position: absolute; inset: 0; width: 100%; height: 100%; opacity: 0; transition: opacity 600ms ease; }
    .subject-media.has-img img { opacity: 1; }
    .subject-media[data-kind="Plant"] img { object-fit: cover; }
    .subject-media[data-kind="Compound"] img { object-fit: contain; padding: 22px; background: #fffdf6; }
    .subject-media .plate-fallback { font-family: var(--display); font-style: italic; font-size: 15px; color: var(--umber); opacity: 0.7; text-align: center; padding: 0 18px; }
    .subject-media.has-img .plate-fallback { display: none; }
    .subject-media.is-loading .plate::after {
      content: ""; position: absolute; inset: 0; z-index: 1;
      background: linear-gradient(100deg, transparent 30%, rgba(255,255,255,0.4) 50%, transparent 70%);
      background-size: 220% 100%; animation: plate-shimmer 1.3s ease-in-out infinite;
    }
    @keyframes plate-shimmer { 0% { background-position: 140% 0; } 100% { background-position: -40% 0; } }
    .subject-media .src-cap { font-family: var(--mono); font-size: 9.5px; letter-spacing: 0.14em; text-transform: uppercase; color: var(--umber); opacity: 0.7; text-align: center; margin-top: 9px; }
    @media (prefers-reduced-motion: reduce) { .subject-media.is-loading .plate::after { animation: none; } }
    @media (max-width: 760px) { .subject-top { flex-direction: column-reverse; gap: 24px; } .subject-media { width: 100%; max-width: 320px; } }
    .subject-type { display: inline-flex; align-items: center; gap: 9px; }
    .subject-type .swatch { width: 11px; height: 11px; border-radius: 50%; display: inline-block; }
    .subject h1 { font-family: var(--display); font-weight: 500; font-style: italic; font-size: 58px; line-height: 1.04; color: var(--forest); margin: 14px 0 0; }
    .subject .common { font-family: var(--display); font-size: 21px; color: var(--umber); margin-top: 6px; }
    .subject .doi { margin-top: 12px; }
    .subject .doi a { font-family: var(--mono); font-size: 13px; letter-spacing: 0.04em; color: var(--umber); border-bottom: 1px solid rgba(122, 90, 58, 0.5); padding-bottom: 2px; transition: color 150ms ease, border-color 150ms ease; }
    .subject .doi a:hover { color: var(--accent); border-color: var(--accent); }
    .subject .doi a .ext { opacity: 0.6; margin-left: 4px; }
    .subject .meta { display: flex; gap: 22px; align-items: baseline; margin-top: 18px; flex-wrap: wrap; }
    .subject .meta .count { font-family: var(--display); font-size: 19px; color: var(--ink); }
    .subject .meta .count b { font-weight: 600; font-variant-numeric: tabular-nums; }
    .graph-link { font-family: var(--display); font-variant-caps: all-small-caps; letter-spacing: 0.1em; font-size: 16px; color: var(--forest); border-bottom: 1px solid var(--forest); padding-bottom: 1px; transition: color 150ms ease, border-color 150ms ease; }
    .graph-link:hover { color: var(--accent); border-color: var(--accent); }
    .dl-link { font-family: var(--display); font-variant-caps: all-small-caps; letter-spacing: 0.1em; font-size: 16px; color: var(--forest); background: none; border: none; border-bottom: 1px solid var(--forest); padding: 0 0 1px; cursor: pointer; transition: color 150ms ease, border-color 150ms ease; }
    .dl-link:hover { color: var(--accent); border-color: var(--accent); }

    /* recorded data properties (datatype attributes) of the focused entity */
    .attrs { margin-top: 30px; border-top: 1px solid var(--rule); }
    .attrs-head { font-family: var(--mono); font-size: 11px; letter-spacing: 0.18em; text-transform: uppercase; color: var(--umber); padding: 16px 0 4px; }
    .attrs-list { display: grid; grid-template-columns: 220px 1fr; gap: 0 28px; }
    .attrs-row { display: contents; }
    .attrs-row .k { font-family: var(--mono); font-size: 12px; letter-spacing: 0.04em; color: var(--umber); padding: 11px 0; border-bottom: 1px dotted rgba(138, 122, 74, 0.4); }
    .attrs-row .v { font-family: var(--display); font-size: 18px; color: var(--ink); padding: 11px 0; border-bottom: 1px dotted rgba(138, 122, 74, 0.4); word-break: break-word; }
    @media (max-width: 760px) {
      .attrs-list { grid-template-columns: 1fr; gap: 0; }
      .attrs-row .k { padding-bottom: 0; border-bottom: none; }
      .attrs-row .v { padding-top: 2px; }
    }

    /* the register itself */
    /* knowledge-graph panel (shared graph.js) */
    .reg-graph { margin: 22px 0 6px; border: 1px solid var(--rule); background: var(--parchment-light); }
    .reg-graph-cap { padding: 13px 22px; border-bottom: 1px solid var(--rule); font-family: var(--display); font-style: italic; font-size: 16px; color: var(--ink); }
    .reg-graph-cap em { color: var(--forest); font-style: italic; }
    .reg-graph-canvas { width: 100%; height: 520px; position: relative; overflow: hidden; }
    @media (max-width: 640px) { .reg-graph-canvas { height: 380px; } }

    .register { padding: 16px 0 64px; }
    .reg-group { margin-top: 40px; }
    .reg-group-head { display: flex; align-items: baseline; gap: 14px; padding-bottom: 8px; border-bottom: 1px solid var(--rule); }
    .reg-group-head .swatch { width: 10px; height: 10px; border-radius: 50%; }
    .reg-group-head .concept { font-family: var(--display); font-style: italic; font-size: 24px; color: var(--forest); }
    .reg-group-head .tally { margin-left: auto; font-family: var(--mono); font-size: 11px; letter-spacing: 0.16em; text-transform: uppercase; color: var(--umber); }

    .reg-row { display: grid; grid-template-columns: 200px 1fr auto; align-items: baseline; gap: 24px; padding: 14px 8px 14px 4px; border-bottom: 1px dotted rgba(138, 122, 74, 0.45); transition: background 120ms ease; }
    .reg-row:hover { background: var(--parchment-light); }
    .reg-row .pred { font-family: var(--mono); font-size: 10px; letter-spacing: 0.14em; text-transform: uppercase; color: var(--umber); padding-top: 4px; }
    .reg-row .pred .arrow { opacity: 0.55; }
    .reg-row .name { font-family: var(--display); font-size: 22px; color: var(--forest); line-height: 1.2; }
    .reg-row .name .latin { font-style: italic; }
    .reg-row .name .vern { font-size: 15px; color: var(--umber); margin-left: 10px; }
    .reg-row .doi-link { display: inline-block; margin-top: 5px; font-family: var(--mono); font-size: 11px; letter-spacing: 0.04em; color: var(--umber); border-bottom: 1px solid rgba(122, 90, 58, 0.4); padding-bottom: 1px; transition: color 150ms ease, border-color 150ms ease; }
    .reg-row .doi-link:hover { color: var(--accent); border-color: var(--accent); }
    .reg-row .doi-link .ext { opacity: 0.6; margin-left: 3px; }
    /* inline multiomics tags (NCBI gene layer) under a target row */
    .reg-row .gtags { display: flex; flex-wrap: wrap; gap: 7px; margin-top: 8px; }
    .gtag { display: inline-flex; align-items: baseline; gap: 6px; font-family: var(--mono); font-size: 11px; letter-spacing: 0.02em; color: var(--ink); background: var(--parchment-deep); border: 1px solid rgba(138, 122, 74, 0.4); padding: 3px 8px; line-height: 1.4; text-decoration: none; transition: background 150ms ease, border-color 150ms ease; }
    a.gtag:hover { background: var(--parchment); border-color: var(--accent); }
    .gtag .gk { font-size: 8.5px; letter-spacing: 0.14em; text-transform: uppercase; color: var(--umber); opacity: 0.85; }
    .gtag.gtag-gene { color: var(--forest); font-weight: 600; }
    .gtag .ext { opacity: 0.55; }
    .reg-row .conn { font-family: var(--mono); font-size: 11px; letter-spacing: 0.08em; color: var(--ink); opacity: 0.6; white-space: nowrap; font-variant-numeric: tabular-nums; }
    .reg-row .conn .dot { display: inline-block; width: 7px; height: 7px; border-radius: 50%; margin-right: 7px; vertical-align: 0.05em; }
    .reg-row a.name-link { display: inline-block; border-bottom: 1px solid transparent; }
    .reg-row:hover a.name-link { border-bottom-color: var(--rule-soft); }

    .reg-empty { padding: 60px 0; text-align: center; color: var(--umber); font-style: italic; font-size: 20px; }

    /* ─── footer (shared) ─── */
    .footer { background: var(--forest-deep); color: var(--parchment); padding: 56px 48px 28px; margin-top: 20px; }
    .footer-inner { max-width: 1180px; margin: 0 auto; }
    .footer-grid { display: grid; grid-template-columns: 2fr 1fr 1fr 1fr; gap: 48px; padding-bottom: 36px; }
    .footer h3 { font-family: var(--display); font-size: 32px; font-style: italic; margin: 0; }
    .footer .tag { font-family: var(--mono); font-size: 10px; letter-spacing: 0.18em; text-transform: uppercase; opacity: 0.6; margin-top: 8px; }
    .footer .blurb { font-size: 15px; line-height: 1.55; opacity: 0.75; margin-top: 18px; max-width: 320px; }
    .footer .col-h { font-family: var(--mono); font-size: 11px; letter-spacing: 0.22em; text-transform: uppercase; opacity: 0.55; margin-bottom: 14px; }
    .footer .col-link { display: block; font-size: 16px; padding: 4px 0; opacity: 0.85; color: inherit; }
    .footer a.col-link:hover { color: var(--accent); opacity: 1; }
    .footer .col-link.active { color: var(--accent); opacity: 1; }
    .footer-bottom a.ft-link { color: inherit; text-decoration: none; border-bottom: 1px solid rgba(243, 233, 205, 0.4); transition: color 150ms ease, border-color 150ms ease; }
    .footer-bottom a.ft-link:hover { color: var(--accent); border-color: var(--accent); }
    .footer-land { font-family: 'EB Garamond', Georgia, serif; font-style: italic; font-size: 14px; line-height: 1.5; opacity: 0.7; padding: 0 0 20px; max-width: 680px; }
    .footer-bottom { border-top: 1px solid rgba(243, 233, 205, 0.2); padding-top: 18px; display: flex; justify-content: space-between; font-family: var(--mono); font-size: 10px; letter-spacing: 0.16em; text-transform: uppercase; opacity: 0.55; }

    @media (max-width: 760px) {
      .wrap { padding: 0 24px; }
      .subject h1 { font-size: 42px; }
      .reg-row { grid-template-columns: 1fr auto; }
      .reg-row .pred { grid-column: 1 / -1; padding-top: 0; }
      .footer-grid { grid-template-columns: 1fr 1fr; gap: 28px; }
    }